1National Key Laboratory of Crop Genetic Improvement and National Center of Plant Gene Research (Wuhan), Huazhong Agricultural University,
Wuhan 430070, China 2Section of Cell and Developmental Biology, University of California, San Diego, La Jolla, CA 92093-0116, USA 3School of Life Sciences, The Chinese University of Hong Kong, Shatin, New Territories, Hong Kong, China 4Guangdong Provincial Key Laboratory of Biotechnology for Plant Development, School of Life Sciences, South China Normal University,
Guangzhou 510631, China
†These authors contributed equally to this article. *Correspondence:
Email: Yunde Zhao (yundezhao@ucsd.edu)