1. State Key Laboratory of Wheat and Maize Crop Science, College of Agronomy and Center for Crop Genome Engineering, Henan Agricultural University, Zhengzhou 450046, China
2. Department of Biological Sciences, University of South Carolina, Columbia, SC 29208, USA *Correspondences: Zheng Qing Fu (zfu@mailbox.sc.edu); Daowen Wang (dwwang@henau.edu.cn)
