1 South China Botanical Garden, the Chinese Academy of Sciences, Guangzhou 510650, China 2 Center of Economic Botany, Core Botanical Gardens, the Chinese Academy of Sciences, Guangzhou 510650, China 3 College of Advanced Agricultural Sciences, University of Chinese Academy of Sciences, Beijing 100049, China
4. Guangdong AIB Polytechnic, Guangzhou 510507, China
† These authors contributed equally to this work. *Correspondence: Xuewu Duan (xwduan@scbg.ac.cn)