1. Department of Plant Pathology and the Ministry of Agriculture Key Laboratory of Pest Monitoring and Green Management, China Agricultural University, Beijing 100193, China; 2. Department of Biochemistry and Biophysics, Texas A&M University, College Station, TX 77843, USA; 3. Department of Horticultural Sciences, Texas A&M University, College Station, TX 77843, USA †These authors contributed equally to this work. *Correspondence: Fuhao Cui (cuifuhao@163.com)