1. Institute of Special Animal and Plant Sciences, Chinese Academy of Agricultural Sciences, Changchun 130000, China; 2. The National Key Facility for Crop Gene Resources and Genetic Improvement, Institute of Crop Sciences, Chinese Academy of Agricultural Sciences, Beijing 100081, China; 3. College of Life Science, Nanjing Agricultural University, Nanjing 210095, China; 4. Agronomy College/National Key Laboratory of Wheat and Maize Crop Science/CIMMYT‐China (Henan) Joint Center of Wheat and Maize, Henan Agricultural University, Zhengzhou 450002, China; 5. Department of Plant Sciences, University of California, Davis, CA 95616, USA; 6. Crop Research Institute, Shandong Academy of Agricultural Sciences, Jinan 250100, China †These authors contributed equally to this work. *Correspondence: Zengyan Zhang (zhangzengyan@caas.cn)