1. The Key Laboratory of Plant Development and Environment Adaptation Biology Ministry of Education, School of Life Science, Shandong University, Qingdao 266237, China; 2. Key Laboratory of Wheat Biology and Genetic Improvement for Low & Middle Yangtze Valley Ministry of Agriculture, Lixiahe Agricultural Institute of Jiangsu Province, Yangzhou 225007, China †These authors contributed equally to this work. *Correspondence: Mengcheng Wang (wangmc@sdu.edu.cn)
