1State Key Laboratory of Crop Stress Adaptation and Improvement, Key Laboratory of Cotton Biology, School of Life Sciences, Henan
University, Kaifeng 475001, China 2State Key Laboratory of Plant Genomics, National Center for Plant Gene Research, Institute of Genetics and Developmental Biology,
Innovation Academy for Seed Design, the Chinese Academy of Sciences, Beijing 100101, China 3Institute of Plant and Microbial Biology, Academia Sinica, Taipei 11529 4Shanghai Center for Plant Stress Biology, Shanghai Institutes for Biological Sciences, the Chinese Academy of Sciences, Shanghai
200032, China 5Department of Horticulture and Landscape Architecture, Purdue University, West LafayetteIndiana 47907, USA
†These authors contributed equally to this work. *Correspondence: Yingfang Zhu (zhuyf@henu.edu.cn)