1 Key Laboratory of Plant Resource Conservation and Germplasm Innovation in Mountainous Region (Ministry of Education), College of
Agricultural Sciences, Guizhou University, Guiyang 550025, China 2 Hubei Engineering Research Center of Viral Vector, Wuhan University of Bioengineering, Wuhan 430415, China 3 National Key Laboratory of Wheat and Maize Crop Science, College of Agronomy, Henan Agricultural University, Zhengzhou 450002,
China