1. College of Agronomy, Qingdao Agricultural University, Qingdao 266109, China; 2. College of Resources and Environment, Qingdao Agricultural University, Qingdao 266109, China; 3. College of Landscape Architecture and Forestry, Qingdao Agricultural University, Qingdao 266109, China; 4. State Key Laboratory of Subtropical Silviculture, School of Forestry and Biotechnology, Zhejiang A&F University, Hangzhou 311300, China †These authors contributed equally to this work. *Correspondences: Gongke Zhou (zhougk@qau.edu.cn); Guohua Chai (chaigh@qau.edu.cn); and Xianfeng Tang (tangxf@qau.edu.cn, Dr. Tang is fully responsible for the distribution of the materials associated with this article)