1. National Key Laboratory for the Development and Utilization of Forest Food Resources, Co‐Innovation Center for Sustainable Forestry in Southern China, State Key Laboratory of Tree Genetics and Breeding, Key Laboratory of State Forestry and Grassland Administration on Subtropical Forest Biodiversity Conservation, College of Life Sciences, Nanjing Forestry University, Nanjing 210037, China

2. Laboratory Center of Life Sciences, College of Life Sciences, Nanjing Agricultural University, Nanjing 210095, China

^†These authors contribute equally to this study.

^*Correspondence: Yanjie Xie (yjxie@njfu.edu.cn; yjxie@njau.edu.cn)