1Development Center of Plant Germplasm Resources, College of Life Sciences,
Shanghai Normal University, Shanghai 200234, China. 2Shanghai Key Laboratory of Plant Molecular Sciences, College of Life Sciences,
Shanghai Normal University, 100 Guilin Road, Shanghai 200234, China. 3Shanghai Center for Plant Stress Biology and Center for Excellence in Molecular Plant
Sciences, Chinese Academy of Sciences, Shanghai 200032, China.
†These authors contributed equally to this work. *Correspondence: Hui Zhang (firstname.lastname@example.org, Dr. Zhang is
fully responsible for the distribution of all materials associated with this
article); Jian-kang Zhu (email@example.com).