1. Frontiers Science Center for Molecular Design Breeding and State Key Laboratory of Plant Environmental Resilience, National Maize Improvement Center, Center for Crop Functional Genomics and Molecular Breeding, Key Laboratory of Biology and Genetic Improvement of Maize (MOA), Beijing Key Laboratory of Crop Genetic Improvement, China Agricultural University, Beijing 100193, China

2. Sanya Institute of China Agricultural University, Sanya 572024, China

^*Correspondence: Feng Tian (ft55@cau.edu.cn)