1. Institute of Advanced Biotechnology, Institute of Homeostatic Medicine, and School of Medicine, Southern University of Science and Technology,Shenzhen 518055, China

2. Shanghai Center for Plant Stress Biology, Center for Excellence in Molecular Plant Sciences, The Chinese Academy of Sciences, Shanghai 201602, China

3. University of Chinese Academy of Sciences, Beijing 100049, China

4. Shandong Shunfeng Biotechnology Co. Ltd., Jinan 250300, China5. National Engineering Laboratory of Crop Stress Resistance Breeding, School of Life Sciences, Anhui Agricultural University, Hefei 230036, China

^†These authors contributed equally to this work.

^*Correspondence: Zhaobo Lang (langzb@sustech.edu.cn)