1. Max Planck Institute of Molecular Plant Physiology, Potsdam‐Golm 14476, Germany

2. Zhejiang Provincial International S&T Cooperation Base for Active Ingredients of Medicinal and Edible Plants and Health, Zhejiang Provincial Key TCM Laboratory for Chinese Resource Innovation and Transformation, School of Pharmaceutical Science, Zhejiang Chinese Medical University, Hangzhou 310053, China

3. School of Breeding and Multiplication (Sanya Institute of Breeding and Multiplication), Hainan University, Sanya 572025, China

4. Yazhouwan National Laboratory, Sanya 572025, China

^*Correspondences: Alisdair R. Fernie (fernie@mpimp-golm.mpg.de, Dr. Fernie is fully responsible for the distribution of all materialsassociated with this article); Guoyin Kai (kaiguoyin@163.com)