1. State Key Laboratory of Biocatalysis and Enzyme Engineering, Hubei Hongshan Laboratory, School of Life Sciences, Hubei University, Wuhan 430062, China; 2. Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences, Shenzhen 518000, China †These authors contributed equally to this work. *Correspondence: Jiang Zhang (zhangjiang@hubu.edu.cn)