1. College of Plant Science and Technology, Huazhong Agricultural University, Wuhan 430070, China; 2. College of Life Science and Technology, Huazhong Agricultural University, Wuhan 430070, China; 3. State Key Laboratory for Development and Utilization of Forest Food Resources, Zhejiang A&F University, Hangzhou 311300, China †These authors contributed equally to this work. *Correspondences: Yuan-Ming Zhang (soyzhang@mail.hzau.edu.cn, Dr. Zhang is fully responsible for the distribution of all materials associated with this article); Jian-Fang Zuo (zuojianfang@zafu.edu.cn)