1. State Key Laboratory of Tropical Crop Breeding, Institute of Tropical Bioscience and Biotechnology, Sanya Research Institute,Chinese Academy of Tropical Agricultural Sciences, Sanya 572024, China

2. National Wheat Engineering Research Center, College of Agronomy, Henan Agricultural University, Zhengzhou 450046, China

3. Department of Biological Sciences, University of South Carolina, Columbia, SC 29208, USA

^*Correspondences: Youxiong Que (queyouxiong@126.com); Zheng Qing Fu (zfu@mailbox.sc.edu, Dr. Fu is fully responsible for thedistribution of all materials associated with this article)