1. The State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, the Chinese Academy of Sciences, Beijing 100101, China 2. University of the Chinese Academy of Sciences, Beijing 100049, China 3. Beijing Institute of Genomics, Chinese Academy of Sciences, National Center for Bioinformation, Beijing 100101, China *Correspondence:Yongbiao Xue (ybxue@genetics.ac.cn)