1State Key Laboratory of Tree Genetics and Breeding, Key Laboratory of Tree
Breeding and Cultivation of the State Forestry Administration, Research Institute
of Forestry, Chinese Academy of Forestry, Beijing 100091, China; 2State Key Laboratory of Subtropical Silviculture, School of Forestry and
Biotechnology, Zhejiang Agriculture & Forestry University, Hangzhou 311300,
China 3Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing
Forestry University, Jiangsu 210037, China; 4Key Laboratory of Agricultural Ecology and Environment, College of Forestry,
Shandong Agricultural University, Tai’an 271018, China; 5School of Agriculture, Ludong University, Yantai 264025, China.