1. College of Horticulture, China Agricultural University, Beijing 100193, China; 2. College of Agriculture and Biotechnology, Zhejiang University, Zijingang Campus, Hangzhou 310058, China; 3. Plant Biology Section, School of Integrative Plant Science, Cornell University, Ithaca 14853, NY, USA; 4. Institute of Horticulture Crops, Xinjiang Academy of Agricultural Sciences, Urumqi 830000, China *Correspondence:Wensuo Jia(jiaws@cau.edu.cn)